Years at Western:
34 (1964-1998)

Subject Specialties:
Entomology, Invertebrate Biology, Marine Biology

Research Interests:
Inbreeding Experiments (Speyeria sp., Papilio Machaon complex), Dwarf Mistletoe insects, biology of Argiope sp. (especially as relates to attracting male moths of the genus Hemileuca), biology of Speyeria Zprene (especially subspecies hippolyta, the Oregon Silverspot, federally listed threatened species), Water beetles of the genus Helophorus (Hydrophilidae)

Current Publications:
Book review - "Swallowtail Butterflies: Their Ecology and Evolutionary Biology" for the Entomological Society of America, 1995.

Photographs - Swallowtail Butterfiles of the Americas, Tyler, Brow and Wilson, Scientific Publishers, 1994.

Book review - "Butterflies of Delmarvn" for the Entomological Society of America, 1994.